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On this page, you can see a list of my publications, and download most of them as a PDF. Your click on any of the links constitutes your request to me for a personal copy of the linked article, and my delivery of a personal copy. Any other use is prohibited. 

Publications by year

2010


Goto, K., Lea, S.E.G., Wills, A.J. and Milton, F. (accepted). Interpreting the effects of image manipulation on picture perception in pigeons (Columba livia) and humans (Homo sapiens). Journal of Comparative Psychology.

The effects of picture manipulations on humans’ and pigeons’ performance were examined in a go/no-go discrimination of two perceptually similar categories, cat and dog faces.  Four types of manipulation were used to modify the images.  Mosaicization and scrambling were used to produce degraded versions of the training stimuli, whilst morphing and cell exchange were used to manipulate the relative contribution of positive and negative training stimuli to test stimuli.  Mosaicization mainly removes information at high spatial frequencies, whereas scrambling removes information at low spatial frequencies to a greater degree.  Morphing leads to complex transformations of the stimuli that are not concentrated at any particular spatial frequency band. Cell exchange preserves high spatial frequency details, but sometimes moves them into the “wrong” stimulus.  The four manipulations also introduce high-frequency noise to differing degrees. Responses to test stimuli indicated that high and low spatial frequency information were both sufficient but not necessary to maintain discrimination performance in both species, but there were also species differences in relative sensitivity to higher and lower spatial frequency information.
Experimental Analysis of Behavior Fellowship, Society for the Advancement of Behavior Analysis
Japan Society for the Promotion of Science
NIMH (MH068426)
EC Framework 6 Project Grant 516542 (NEST) 


Pothos, E.M. and Wills, A.J. (Eds.) (in press). Formal approaches in categroization. Cambridge University Press.

Includes chapters by Nosofsky, J.D. Smith, Ashby, McClelland,  Kruschke, McLaren, Tenenbaum, Chater, Gureckis, Langley, Rehder, Murphy, Medin, Caramazza, and co-authors.

Beesley, T., Wills, A.J. and Le Pelley, M.E. (2010). Syntactic transfer in artificial grammar learning. Psychonomic Bulletin & Review, 17, 1, 122-128.

In an artificial grammar learning experiment participants were trained with instances of one grammatical structure, before completing a test phase in which they were required to discriminate grammatical from randomly created strings. Importantly, the underlying structure used to generate test strings was different from that used to generate the training strings. Despite the fact that grammatical training strings were more similar to non-grammatical test strings than grammatical test strings, this manipulation resulted in a positive transfer effect, compared to controls trained with non-grammatical strings. It is suggested that training with grammatical strings leads to an appreciation of set variance which aids the detection of grammatical test strings in AGL tasks. The analysis presented demonstrates that it is useful to conceptualise test performance in AGL as a form of unsupervised category learning. 

ESRC Grant RES-000-22-1779
EC Framework 6 Project Grant 516542 (NEST)

Hopewell, L., Leaver, L.A., Lea, S.E.G., and Wills, A.J. (2010). Grey squirrels (Sciurus carolinensis) show a feature-negative effect specific to social learning. Animal Cognition 13, 2, 219-227.

Previous laboratory studies on social learning suggest that some animals can learn more readily if they observe a conspecific demonstrator perform the task unsuccessfully and so fail to obtain a food reward than if they observe a successful demonstrator that obtains the food. This effect may indicate a difference in how easily animals are able to associate different outcomes with the conspecific or could simply be the result of having food present in only some of the demonstrations. To investigate we tested a scatter-hoarding mammal, the eastern grey squirrel, on its ability to learn to choose between two pots of food after watching a conspecific remove a nut from one of them on every trial. Squirrels that were rewarded for choosing the opposite pot to the conspecific chose correctly more frequently than squirrels rewarded for choosing the same pot (a feature-negative effect). Another group of squirrels was tested on their ability to choose between the two pots when the rewarded option was indicated by a piece of card. This time, squirrels showed no significant difference in their ability to learn to choose the same or the opposite pot. The results add to anecdotal reports that grey squirrels can learn by observing a conspecific and suggest that even when all subjects are provided with demonstrations with the same content, not all learning occurs equally. Prior experience or expectations of the association between a cue (a conspecific) and food influences what can be learned through observation whilst previously unfamiliar cues (the card) can be associated more readily with any outcome.

EC Framework 6 Project Grant 516542 (NEST)

2009


Wills, A.J., Lea, S.E.G., Leaver, L.A., Osthaus, B., Ryan, C.M.E., Suret, M.B., Bryant, C.M.L., Chapman, S.J.A. and Millar, L. (2009). A Comparative Analysis of the Categorization of Multidimensional Stimuli: I. Unidimensional Classification Does not Necessarily Imply Analytic Processing; Evidence From Pigeons (Columba livia), Squirrels (Scurius carolinensis), and Humans (Homo sapiens). Journal of Comparative Psychology, 123, 4, 391-405.

Pigeons, gray squirrels and undergraduates learned discrimination tasks involving multiple mutually redundant dimensions.  There were two sets of experiments.  Within each set, stimuli and procedures were as closely similar for the different species as possible, but they differed sharply between sets.  First, pigeons and undergraduates learned conditional discriminations between stimuli composed of three spatially separated dimensions, after first being trained to discriminate the individual elements of the stimuli.  They were then tested with stimuli in which one of the three dimensions took an anomalous value.  The majority of both species categorized test stimuli by their overall similarity to training stimuli, but some individuals of both species categorized them according to a single dimension.  Secondly, squirrels, pigeons and undergraduates learned go/no-go discriminations using multiple simultaneous presentations of stimuli composed of three spatially integrated, highly salient dimensions.  In tests, the tendency to categorize stimuli including anomalous dimension values unidimensionally was higher than in the first set of experiments and did not differ significantly between species.  We conclude that unidimensional categorization of multidimensional stimuli is not diagnostic for analytic cognitive processing, and that any differences in behavior between humans and pigeons in such tasks are not due to special features of avian visual cognition.  

ISSN: 0735-7036

EC Framework 6 Project Grant 516542 (NEST)


Lea, S.E.G., Wills, A.J., Leaver, L.A., Ryan, C.M.E., Bryant, C.M.L. and Millar, L. (2009). A Comparative analysis of the categorization of multidimensional stimuli: II. Strategic information search in humans (Homo sapiens) but not in pigeons (Columba livia). Journal of Comparative Psychology, 123, 4, 406-420.

Pigeons and undergraduates learned conditional discriminations involving multiple spatially separated stimulus dimensions.  Under some conditions the dimensions were made available sequentially.  In three experiments the dimensions were all perfectly valid predictors of the response that would be reinforced and mutually redundant; in two others they varied in validity.  In tests with stimuli where one of the three dimensions took an anomalous value, most but not all individuals of both species categorized them in terms of single dimensions.  When information was delivered as a function of the passage of time, some students but no pigeons waited for the most useful information, especially when the cues differed in objective validity.  When the subjects could control information delivery, both species obtained information selectively.  When cue validities varied, almost all students tended to choose the most valid cues, and when all cues were valid, some chose the cues by which they classified test stimuli.  Only a few pigeons chose the most useful information in either situation.  Despite their tendency to unidimensional categorization, the pigeons showed no evidence of rule-governed behavior, but students followed a simple Take the Best rule.

ISSN: 0735-7036

ESRC Grant RES-000-22-1779
EC Framework 6 Project Grant 516542 (NEST)

Milton, F., Wills, A.J. and Hodgson, T.L. (2009). The neural basis of overall similarity and single-dimension sorting. Neuroimage, 46, 319-326.

The ability to group stimuli into meaningful categories is fundamental to natural behavior. Raw perceptions would be useless without an ability to classify items as, for example, threat or food. Previous work suggests that people have a tendency to group stimuli either on the basis of a single dimension or by overall similarity (e.g., Milton, Longmore, & Wills, 2008). It has recently been suggested that overall similarity sorting can engage similar rule-based processes to single-dimension sorting and, in addition, requires greater use of working memory (Milton & Wills, 2004). These predictions were tested in an event-related fMRI study of spontaneous categorization. Results showed a striking overlap of activation between overall similarity and single-dimension sorting indicating engagement of common neural processes.  Furthermore, overall similarity sorting recruited additional activity in bilateral precuneus, right cuneus, left cerebellum, left postcentral gyrus, right thalamus and right ventrolateral frontal cortex (VLFC). Our findings suggest that overall similarity sorting can be the result of rule-based processes and highlight a potential role for right VLFC in integrating multi-dimensional sensory information to form conceptual categories.

ESRC Grants PTA-030-2003-00287; PTA-026-27-1256; RES-000-22-1779
Great Western Research Fellowship (GWR-34)
EC Framework 6 project grant 516542 (NEST)

Milton, F. and Wills, A.J. (2009). Long-term persistence of sort strategy in free classification. Acta Psychologica. 130, 161-167.

Two free classification experiments that investigate the persistence of sort strategy are reported. Participants tend to persist with their initial categorization type (family resemblance or unidimensional) for the remaining sorts, overriding the effects of otherwise influential stimulus properties. Sort type was found to persist even after a one-week delay. Stimulus-driven models of free classification (e.g., the SUSTAIN model, Love, Medin & Gureckis, 2004) cannot predict the sort type persistence effects we observe, but they are naturally accounted for by theories that posit strategic selection of a problem-solving strategy (e.g., Hypothesis theory, Levine, 1971).

ESRC Grants PTA-030-2003-00287; PTA-026-27-1256; RES-000-22-1779
Great Western Research Fellowship (GWR-34)
EC Framework 6 project grant 516542 (NEST)

Wills, A.J. (2009).  Prediction Errors and Attention in the Presence and Absence of Feedback. Current Directions in Psychological Science, 18, 95-100.

Contemporary theories of learning typically assume that learning is driven by prediction errors—in other words, that we learn more when our predictions turn out to be incorrect than we do when our predictions are correct. Results from the recording of electrical brain activity suggest one mechanism by which this might happen; we seem to direct visual attention toward the likely causes of previous prediction errors. This can happen very rapidly—within less than 200 milliseconds of the error-causing object being presented. It is tempting to infer that if learning is driven by prediction errors then little can be learned in the absence of feedback. Such a conclusion is unwarranted. In fact, the substantial learning that is sometimes the result of simple exposure to objects can also be explained by processes of directing attention toward the likely causes of previous prediction errors.

ISSN: 0963-7214

BBSRC Grant 9/S17109
EC Framework 6 project grant 516542 (NEST)

Milton, F. and Wills, A.J. (2009).  Eye movements in overall similarity and single-dimension sorting. In N.A. Taatgen & H. van Rijn (Eds.), Proceedings of the Thirty-First Annual Conference of the Cognitive Science Society (pp. 1512-1517). Austin, TX: Cognitive Science Society.

A free classification study is presented that uses eye-tracking to better characterize the strategies that are employed in the creation of overall similarity and single-dimension categories. The number of fixations across the dimensions and the proportion of dimensions fixated were significantly greater for overall similarity sorters than for single-dimension sorters. Single-dimension sorters generally fixated a single dimension from the outset, a strategy in line with the principles of SUSTAIN (Love, Medin, & Gureckis, 2004). The pattern of eye movements is consistent with the idea that overall similarity sorting can be a time consuming process that involves greater perceptual processing of the stimuli than single-dimension sorting.

ESRC Grants PTA-030-2003-00287; PTA-026-27-1256; RES-000-22-1779
EC Framework 6 project grant 516542 (NEST)

2008 


Milton, F., Longmore, C.A. and Wills, A.J. (2008). Processes of overall similarity sorting in free classification. Journal of Experimental Psychology: Human Perception and Performance, 34, 676-692.

The processes of overall similarity sorting were investigated in five free classification experiments. Experiments 1 and 2 demonstrate that increasing time pressure can reduce the likelihood of overall similarity categorization. Experiment 3 showed a concurrent load also reduced overall similarity sorting. These findings suggest overall similarity sorting can be a time consuming, analytic process. Such results appear contrary to the idea that overall similarity is a non-analytic process (e.g. Ward, 1983), but are in line with Milton and Wills’s (2004) dimensional summation hypothesis, and the stochastic sampling assumptions of EGCM (Lamberts, 2000). Experiments 4 and 5 demonstrate that the relationship between stimulus presentation time and overall similarity sorting is non-monotonic, and the shape of the function is consistent with the idea that the three aforementioned processes operate over different parts of the time course

BBSRC Grant 9/S17109
ESRC Grants PTA-030-2003-00287; PTA-026-27-1256; RES-000-22-1779
EC Framework 6 project grant 516542 (NEST)

Lea, S.E.G. and Wills, A.J. (2008). Use of multiple dimensions in learned discriminations, Comparative Cognition and Behavior Reviews, 3, 115-133.

Many naturally-occurring categories vary across multiple stimulus dimensions (e.g. size, color, texture). Where humans categorize multidimensional stimuli on the basis of a single dimension this has classically been taken to indicate use of a rule that could be verbalized, whilst when they sort on the basis of all the stimulus dimensions (known as 'overall similarity' or 'family resemblance' sorting) it has been taken to indicate a more basic, implicit, automatic process (sometimes referred to as associative). However, a review of the literature on concept discrimination in pigeons suggests that animals often discriminate on the basis of one dominant dimension; and recent human experiments indicate that situations particularly conducive to more complex cognitive processes can actually increase family resemblance sorting in humans. Are we to conclude from these two counter-examples that pigeons can use rules and that humans, when in conditions conducive to complex thought, choose to respond associatively? In an effort to resolve this apparent paradox, we are conducting experiments in which humans and pigeons are exposed to multidimensional category discrimination tasks under closely similar conditions. The results throw light on what it means for behavior to be governed by a rule, and suggest that there is no evidence that even a non-verbal rule can be said to be involved in pigeons’ choices in these conditions, despite the fact that under some conditions a single dimension may dominate their behavior.

ESRC Grant RES-000-22-1779
EC Framework 6 Project Grant 516542 (NEST)

Milton, F. and Wills, A.J. (2008). The influence of perceptual difficulty on family resemblance sorting. Proceedings of the Thirtieth Annual Conference of the Cognitive Science Society, 2273 - 2278.

The impact of perceptual difficulty on the prevalence of family resemblance sorting was investigated in a free classification experiment. There were two between-subject conditions, high perceptual difficulty and low perceptual difficulty and participants were asked to sort the stimuli into two groups in the way that seemed most appropriate to them. The results showed that participants in the low perceptual difficulty condition sorted by family resemblance to a greater extent than participants in the high perceptual difficulty condition. This finding extends the work of Milton and Wills (2004), who showed that the level of spatial integration in stimuli is an important determinant of sorting behavior, but whose results were inconclusive on the issue of perceptual difficulty. The current experiment adds to the growing body of work which demonstrates that free sorting behavior can be influenced in a variety of different ways.
BBSRC Grant 9/S17109
ESRC Grants PTA-030-2003-00287; PTA-026-27-1256; RES-000-22-1779
EC Framework 6 project grant 516542 (NEST)

2007


Haslam, C.,Wills, A.J., Haslam, S.A., Kay, J., Baron, R. and McNab, F. (2007). Does maintenance of colour categories rely on language? Evidence to the contrary from a case of semantic dementia. Brain and Language, 103, 251–263.

Recent neuropsychological evidence, supporting a strong version of Whorfian principles of linguistic relativity, has reinvigorated debate about the role of language in colour categorization.  This paper questions the methodology used in this research and uses a novel approach to examine the unique contribution of language to categorization behaviour.  Results of three investigations are reported.  The first required development of objective measures of category coherence and consistency to clarify questions about healthy control performance on the freesorting colour categorization task used in previous studies. Between-participant consistency was found to be only moderate and the number of colour categories generated was found to vary markedly between individuals.  The second study involved longitudinal neuropsychological examination of a patient whose colour categorization strategy was monitored in the context of a progressive decline in language due to semantic dementia.  Performance on measures of category coherence and consistency was found to be relatively stable over time despite a profound decline in the patient’s colour language.  In a final investigation we demonstrated that, for both the patient and controls, between- and within-participant consistency were higher than expected by (a) random sorting and (b) sorting perceptually similar chips together.   These findings indicate that the maintenance of colour categorization need not depend on language.

ISSN: 0093-934X
Imprint: ELSEVIER

BBSRC Grant 9/S17109

ESRC Grant RES-000-22-1779

Wills, A.J., Lavric, A, Croft, G. and Hodgson, T.L (2007). Predictive learning, prediction errors and attention: Evidence from event-related potentials and eye-tracking. Journal of Cognitive Neuroscience. 19, 843-854.

Prediction error (“surprise”) affects the rate of learning: we learn more rapidly about cues for which we initially make incorrect predictions than cues for which our initial predictions are correct.  The current studies employ electrophysiological measures to reveal early attentional differentiation of events that differ in their previous involvement in errors of predictive judgment. Error-related events attract more attention, as evidenced by features of event-related scalp potentials previously implicated in selective visual attention (selection negativity, augmented anterior N1). The earliest differences detected occurred around 120 ms after stimulus onset, and  distributed source localization (LORETA) indicated that the inferior temporal regions were one source of the earliest differences. In addition, stimuli associated with the production of prediction errors show higher dwell times in an eye-tracking procedure. Our data support the view that early attentional processes play a role in human associative learning.
BBSRC Grant 9/S17109
EC Framework 6 project grant 516542 (NEST)

2006


Wills, A.J., Noury, M., Moberly, N.J. and Newport, M. (2006). Formation of category representations. Memory & Cognition, 34, 17-27. 

Many formal models of categorization assume, implicitly or explicitly, that categorization results in the formation of direct associations from representations of the presented stimuli to representations of the experimentally-provided category labels. In three categorization experiments employing a polymorphous classification structure (Dennis, Hampton, & Lea, 1973) and a partial reversal, “optional shift” procedure, (Kendler, Kendler, & Wells, 1960) we provide evidence consistent with the hypothesis that learning a new classification problem results in the creation of category representations that mediate between representations of stimulus and label. This hypothesis can be instantiated through the AMBRY model (Kruschke, 1996).
BBSRC Grant 9/S17109
EC Framework 6 project grant 516542 (NEST)

Lea, S.E.G., Wills, A.J. and Ryan, C.M.E. (2006). Why are artificial polymorphous concepts so hard for birds to learn? Quarterly Journal of Experimental Psychology, 59, 251-267.

Stimulus sets defined in terms of artificial polymorphous concepts have frequently been used in experiments to investigate the mechanisms of discrimination of natural concepts, both in humans and in other animals.  However such stimulus sets are frequently difficult for either animals or humans to discriminate.  Properties of artificial polymorphous stimulus sets that might explain this difficulty include the complexity of the individual stimuli, the unreliable reinforcement of individual positive features, attentional load, difficulties in discriminating some stimulus dimensions, memory load, and a lack of the correlation between features that characterises natural concepts.  An experiment using chickens as subjects and complex artificial visual stimulus sets investigated these hypotheses by training the birds in discriminations that were not polymorphous but did have some of the properties listed above.  Discriminations that involved unreliable reinforcement or high attentional load were found to approach the difficulty of polymorphous concept discriminations, and these two factors together were sufficient to account for the entire difficulty.  The usual kind of artificial polymorphous concept may not be a good model for natural concepts as they are perceived and discriminated by birds.  A RULEX account of natural concept learning may be preferable.

BBSRC Grant 9/S17109

2005


Le Pelley, M.E., Oakeshott, S.M, Wills, A.J. and McLaren, I.P.L. (2005). The outcome-specificity of learned predictiveness effects: Parallels between human causal learning and animal conditioning. Journal of Experimental Psychology: Animal Behavior Processes, 31, 226-236.

Two experiments examined the outcome-specificity of a learned predictiveness effect in human causal learning. Experiment 1 indicated that prior experience of a cue–outcome relationship modulates learning about that cue with respect to a different outcome from the same affective class, but not to an outcome from a different affective class. Experiment 2 ruled out an interpretation of this effect in terms of context-specificity. These results indicate that learned predictiveness effects in human causal learning index an associability that is specific to a particular class of outcomes. Moreover they mirror demonstrations of the reinforcer-specificity of analogous effects in animal conditioning, supporting the suggestion that, under some circumstances, human causal learning and animal conditioning reflect the operation of common associative mechanisms.

ESRC grant RG36281

Wills, A.J. (2005). New directions in human associative learning. Lawrence Erlbaum Associates.

The editor and authors of this book have forged a new synthesis of the work on Human Associative Learning from Pavlov through the 21st century. It is divided into three sections: an introduction to the recent data and controversies in the study of associative learning; recent developments in the formal theories of how associative learning occurs, and applied work on human associative learning, particularly its application to depression and to the development of preferences. This book is designed to be accessible to undergraduates, providing a clear illustration of the principles of animal cognition and the contemporary study of human cognition.


Wills, A.J. (2005). Association and cognition. In A.J.Wills (Ed.). New directions in human associative learning. Lawrence Erlbaum Associates. pp. 1-9.

    In April 1903, Pavlov presented his early work on the conditioned reflex to the International Congress of Medicine, Madrid (Pavlov, 1903/1928a). Two of the, no doubt numerous, events that celebrated the centenary of this event were the Spring meeting of the Experimental Psychology Society at Exeter (for which I was the local organizer) and the annual Associative Learning Symposium organized by Professors Rob Honey and John Pearce of Cardiff University.
    As the date of the Experimental Psychology Society (EPS) conference approached, my thoughts turned increasingly to the nature of the man and the work we were about to celebrate, and how both were considered today. I repeatedly recalled a conversation with a colleague about a year earlier that started with him saying, 'Associative learning ... isn't that something they did in the 1950s when they thought pigeons could be trained to guide missiles?"
    It's probably best that this particular opinion remains anonymous but, in less extreme forms, it probably reflects the attitude of a considerable minority of today's professional psychologists. Breaking down the opinion into its constituents, it seems to center on the ideas that the study of associative learning is (a) entirely concerned with nonhuman animals, and is (b) part of an episode in the history of psychology that ended with the "cognitive revolution" of the 1960s and 1970s. Both components of this opinion are fundamentally incorrect, as I indicate later in this chapter. However, first there is a need to define terms....

Wills, A.J. (2005). Connectionist models of human associative learning. In A.J.Wills (Ed.). New directions in human associative learning. Lawrence Erlbaum Associates. pp. 95-99.

In chapter 1,1 drew a distinction between two quite different concepts that sometimes both attract the term associative learning. On some occasions, "associative learning" is used to define a particular type of problem that an organism has to solve. On other occasions, "associative learning" is used as a theoretical statement about the sorts of mental processes by which the organism solves this type of problem...

Zwickel, J. and Wills, A.J. (2005). Integrating associative models of supervised and unsupervised categorization. In A.J.Wills (Ed.). New directions in human associative learning. Lawrence Erlbaum Associates. pp. 101-123.

In this chapter, we describe three basic associative theories of learning as they might be applied to the problem of category learning. First, we describe Hebbian learning, which was one of the earliest formal associative theories of learning. Second, we describe competitive learning, which can, in some ways, been seen as a development of Hebbian learning, and is a theory of unsupervised learning. next, we describe the Rescorla-Wagner theory, which is a theory of supervised learning. We then argue that there is a need for a mechanism that can use feedback when it is available (supervised learning) but will continue to learn in feedback is absent (unsupervised learning). We propose a possible mechanism that involves adding certain aspects of Rescorla-Wagner theory to competitive learning. Our proposed system makes a claer prediction about people's behavior in a free-classification task, and we describe how we have begun to test that prediction. The chapter ends with a consideration of other theoretical approaches to the problem of category learning, both within the domain of associative learning and more generally.


Wills, A.J. (2005). Applications and extension. In A.J.Wills (Ed.). New directions in human associative learning. Lawrence Erlbaum Associates. pp. 189-191.

In the final two chapters of this book, we turn to phenomena whose practical implications are more direct than some of the phenomena previously discussed....

2004


Wills, A.J., Suret, M.B. and McLaren, I.P.L. (2004). The role of category structure in determining the effects of stimulus preexposure on categorization accuracy. Quarterly Journal of Experimental Psychology, 57B, 79-88.

What are the effects of pre-exposure of stimuli on participants' subsequent ability to categorize them accurately? An experiment employing artificial, abstract, visual stimuli confirms that, for adult humans, the effect of pre-exposure is dependent upon category structure. Whether pre-exposure has beneficial or detrimental effects is shown to be dependent on the way category examples are generated from the category base patterns. The results are predicted by salience reduction accounts of perceptual learning but may be problematic for stimulus differentiation accounts.

Junior Research Fellowship, Emmanuel College, Cambridge
BBSRC Grant 9/S17109

Milton, F. and Wills, A.J. (2004). The influence of stimulus properties on category construction. Journal of Experimental Psychology. Learning, Memory, and Cognition. 30, 407-415.

It has been demonstrated that when individuals free classify stimuli presented simultaneously in an array, they have a preference to categorize by a single dimension (e.g. Medin, Wattenmaker & Hampson, 1987). However, when people are encouraged to categorize items sequentially, initial research has suggested that people sort by “family resemblance” – grouping by overall similarity (Regehr &  Brooks, 1995). Our studies extended this research, producing three main findings. First, the sequential procedure introduced by Regehr & Brooks (1995) does not always produce a preference for family resemblance sorts. Second, sort strategy in a sequential procedure is highly sensitive to subtle variations in stimulus properties. Third, and surprisingly in the context of previous research, spatially separable stimuli evoked more family resemblance sorts than stimuli of greater spatial integration. It is suggested that the family resemblance sorting observed is the result of an analytic strategy as opposed to a perceptually driven non-analytic strategy.

ESRC Grant PTA-030-2003-00287
BBSRC Grant 9/S17109

Goto, K, Wills, A.J. and Lea, S.E.G. (2004). Global-feature classification can be acquired more rapidly than local-feature classification in both humans and pigeons. Animal Cognition. 7, 109-113.

When humans process visual stimuli, global information often takes precedence over local information. In contrast, some recent studies have pointed to a local precedence effect in both pigeons and nonhuman primates. In the experiment reported here, we compared the speed of acquisition of two different categorizations of the same four geometric figures. One categorization was on the basis of a local feature, the other on the basis of a readily apparent global feature. For both humans and pigeons, the global-feature categorization was acquired more rapidly. This result reinforces the conclusion that local information does not always take precedence over global information in nonhuman animals.

BBSRC Grant 9/S17109

Bryant, C.M., Jones, G.J.F., and Wills, A.J. (2004). Integration of psychological models in the design of artificial creatures. Proceedings of the AISB 2004 symposium on Emotion, Cognition and Affective Computing. pp. 9-20 (ISBN: 1 902956 36 6).

Artificial creatures form an increasingly important component of interactive computer games. Examples of such creatures exist which can interact with each other and the game player and learn from their experiences. However, we argue, the design of the underlying architecture and algorithms has to a large extent overlooked knowledge from psychology and cognitive sciences. We explore the integration of observations from studies of motivational systems and emotional behaviour into the design of artificial creatures. An initial implementation of our ideas using the “sim agent” toolkit illustrates that physiological models can be used as the basis for creatures with animal like behaviour attributes. The current aim of this research is to increase the “realism” of artificial creatures in interactive game-play, but it may have wider implications for the development of AI.

BBSRC Grant 9/S17109

2003


Lochmann, T. & Wills, A.J. (2003). Predictive history in an allergy prediction task. In Schmalhofer, F., Young, R. M. & Katz, G. (Eds.) Proceedings of EuroCogSci 03: The European Cognitive Science Congerence. Mahwah, New Jersey: Lawrence Erlbaum Associates. pp. 217-222.

Two experiments are reported that demonstrate rate of learning in an allergy prediction task can be affected by the predictive history of the cues involved, even if that history relates to outcomes different to those being currently learned about. Predictive history is defined here as a cue’s prior status as either a good or a poor predictor of outcomes. Our results are problematic for commonly employed associative theories of human contingency learning but also provide evidence for the sort of associability-change process envisaged by the Mackintosh (1975) theory.

BBSRC Grant 9/S17109

Welham, A., Schnadt, M. and Wills, A.J. (2003). Speeded categorization: the effects of perceptual processing and decision-making time. Proceedings of the 25th Annual Conference of the Cognitive Science Society. pp. 1218-1223.

The effects of limited processing time were investigated for the binary categorization of artificial multidimensional objects. Following Lamberts and Freeman (1999), in the first stage of the experiment participants learnt to categorize 9 stimuli into two categories. In the second stage, the same stimuli were presented for categorization, and both the display time, and the time available in which to make a decision, were varied independently. It was found that each of these variables had a significant effect on accuracy of categorization, as well as response latency. Lamberts and Freeman (1999) demonstrated that restricting presentation time of a certain stimulus in their category structure caused a reversal in category assignment. We found evidence of the same reversal, but it was dependent on the time available to make a decision rather than the duration of stimulus display. Importantly, changes in accuracy due to response deadline were not explicable in terms of truncation of processing by the limited time. The study provides an empirical investigation of the intuitive notion that both perceptual processing and decision making components are time dependent.

BBSRC Grant 9/S17109

2002


Zwickel, J. and Wills, A.J. (2002). Is competitive learning an adequate account of free classification? Proceedings of the 24th Annual Conference of the Cognitive Science Society. pp. 982-987.

Rumelhart & Zipser's (1986) competitive learning algorithm is an account of unsupcrvised learning and, as such, might be considered a potential model of free classification behavior in humans. However, selective learning effects (e.g. Dickinson, Shanks & Evenden, 1984) suggest that human learning, ai least under conditions of feedback, may be better characterized by an error-correcting system. An experiment is reported that provides preliminary evidence for the existence of a selective learning effect in free classification. Simulations indicate that Rumelhart & Zipser's algorithm does not provide an adequate account of the behavior observed, whilst an error-correcting variant of competitive learning does.

Research Fund, University of Exeter.

Wills, A.J. (2002). Adapting to a response deadline in categorization. Proceedings of the 24th Annual Conference of the Cognitive Science Society. 938-943.

The effect of a response deadline on categorical decisions was investigated. Time available for response was manipulated in the test phase, along with stimulus difficulty. Effects of these manipulations were observed in response accuracy, and in the mean, standard deviation and skew of the reaction times. The effects observed demonstrate that participants responded to the deadline in an adaptive manner - reducing their reaction time to long-latency decisions whilst leaving short latency decisions relatively unaffected. A simple connectionist model of categorical decisions (Wills & McLaren, 1997) is shown to account for this behavior.

ESRC Grant
Junior Research Fellowship, Emmanuel College, Cambridge 

Wills, A.J. (2002). Representing information. Psychology. Vol 3: Thinking and knowing. Grolier.

Wills, A.J. (2002).  Behaviorism. Psychology. Vol. 1: History of psychology. Grolier.

2000


Wills, A.J., Reimers, S., Stewart, N., Suret, M. and  McLaren, I.P.L. (2000). Tests of the ratio rule in categorization. Quarterly Journal of Experimental Psychology. 53A, 983-1011.

Many theories of learning and memory (e.g., connectionist, associative, rational, exemplar based) produce psychological magnitude terms as output (i.e., numbers representing the momentary level of some subjective property). Many theories assume that these numbers may be translated into choice probabilities via the ratio rule, also known as the choice axiom (Luce, 1959) or the constant-ratio rule (Clarke, 1957).We present two categorization experimentsemploying artificial, visual, prototype-structured stimuli constructed from12 symbols positioned on a grid. The ratio rule is shown to be incorrect for these experiments, given the assumption that the magnitude terms for each category are univariate functions of the number of category-appropriate symbols contained in the presented stimulus. A connectionist winner-take-all model of categorical decision (Wills & McLaren, 1997) is shown to account for our data given the same assumption. The central feature underlying the success of this model is the assumption that categorical decisions are based on a Thurstonian choice process (Thurstone, 1927, Case V) whose noise distribution is not double exponential in form.

BBSRC Grant
ESRC Grant

1998


Wills, A.J. and McLaren, I.P.L. (1998). Perceptual learning and free classification. Quarterly Journal of Experimental Psychology. 51B, 235-70.

Two experiments are reported that investigate the effects of stimulus preexposure on discrimination performance in a free classification task, using adult humans as subjects. In free classification subjects are asked to put stimuli into gruops in any way that seems reasonable or sensible to them. Experiment 1 shows that the effect of preexposure is contingent on stimulus structure. Experiment 1b is the first demonstration of a retardation in learning as a consequence of simple preexposure in adult human subjects (previous demonstrations have relied on incidental or masked preexposure). Experiment 2 further supports the conclusions of Experiment 1 and extends them with the demonstration that stimulus similarity is a crucial factor. Taken together, these experiments rule out a class of attention-based explanations of the phenomena reported here. The experiments also provide novel information about the effects of preexposure. Preexposure can change the actual classifications subjects form in addition to altering the rate at which they are formed. Implications of these results for current theories of category formation and perceptual learning are considered.

BBSRC Grant
BBSRC Studentship


Jones, F.W., Wills, A.J. and McLaren, I.P.L. (1998). Perceptual categorization: connectionist modelling and decision rules. Quarterly Journal of Experimental Psychology. 51B, 33-58.

Although it is currently popular to model human associative learning using connectionist networks, the mechanism by which their output activations are converted to probabilities of response has received relatively little attention. Several possible models of this decision process are considered here, including a simple ratio rule, a simple difference rule, their exponential versions, and a winner-take-all network. Two categorization experiments that attempt to dissociate these models are reported. Analogues of the experiments were presented to a single-layer, feed-forward, delta-rule network. Only the exponential ratio rule and the winner-take-all architecture, acting on the networks’ output activations that corresponded to responses available on test, were capable of fully predicting the mean response results. In addition, unlike the exponential ratio rule, the winner-take-all model has the potential to predict latencies. Further studies will be required to determine whether latencies produced under more stringent conditions conform to the model’ s predictions.

BBSRC Grant

1997


Wills, A.J. and McLaren, I.P.L. (1997). Generalization in human category learning: A connectionist explanation of differences in gradient after discriminative and non-discriminative training. Quarterly Journal of Experimental Psychology. 50A, 607-630.
Two experiments are reported that investigate the difference in g radient of generalization observed between one-category (non-discriminative) and two-category (discriminative) training. Extrapolating from the results of a number of animal lear ning studies, it was predicted that the gradient should be steeper under discriminative tr aining. The first experiment concerns this basic prediction for the stimuli used, which were novel, prototype-structured, and constructed from 12 symbols positioned on a grid. An explanation for the effect, based on the Rescorla-Wagner theory of Pavlovian conditioning (Rescorla & Wagner, 1972), is that under non-discriminative tr aining "incidental stimuli" have significant control over responding, whereas under discriminative training they do not. Incidental stimuli are those aspects of the stimulus, or the surrounding context, that are not differentially reinforced under discriminative training. This explanation leads to the prediction that a comparable effect of blocked ver sus intermixed discriminative training should also be found. This prediction is disconfirmed by the second experiment. An alternative model, still based on the Rescorla-Wagner theory but with the addition of a decision mechanism comprising a threshold unit and a competitive network system, is proposed, and its ability to predict both the choice probabilities and the pattern of response times found is evaluated via simulation.

BBSRC Grant